Cicadas (Hemiptera: Cicadidae) from Samar and Leyte, Philippines, with a new species


Lee, Y.J.

Journal of Asia-Pacific Entomology 14(3): 253-257

2011


Contents
lists
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at
ScienceDirect
Journal
of
Asia-Pacific
Entomology
journal
homepage:
www.elsevier.com/locate/jape
14
of
21,La-Pacific
E111.11101.11gy
Cicadas
(Hemiptera:
Cicadidae)
from
Samar
and
Leyte,
Philippines,
with
a
new
species
Young
June
Lee
*
Department
of
Ecology
and
Evolutionary
Biology,
University
of
Connecticut,
75
North
Eagleville
Road,
Storrs,
CT
06269,
USA
ARTICLE INFO
ABSTRACT
Journal
of
Asia-Pacific
Entomology
14
(2011)
253-257
Article
history:
Received
2
December
2010
Revised
11
January
2011
Accepted
12
January
2011
Available
online
20
January
2011
Keywords:
Taxonomy
Fauna
Checklist
Distribution
Huechys
funebris
This
paper
provides
the
first
faunal
checklist
for
Cicadidae
(Hemiptera)
from
Samar
and
Leyte,
Philippines,
comprising
13
species
belonging
to
7
genera.
A
new
species,
Huechys
funebris
sp.
nov.,
is
described
from
Leyte.
Oncotympana
pallidiventris
Stal,
Dundubia
vaginata
(Fabricius),
and
Nelcyndana
sp.
are
newly
added
to
the
cicada
fauna
of
Samar.
Information
on
geographic
distributions
is
provided.
Korean
Society
of
Applied
Entomology,
Taiwan
Entomological
Society
and
Malaysian
Plant
Protection
Society,
2011.
Published
by
Elsevier
B.V.
All
rights
reserved.
Introduction
This
paper
contributes
to
the
knowledge
of
cicada
diversity
in
Samar
and
Leyte,
providing
their
first
faunal
checklist.
It
is
the
fifth
in
a
series
of
faunal
studies
on
the
cicadas
of
the
major
Philippine
islands
after
those
of
Palawan,
Panay,
Mindanao,
and
Luzon
(Lee,
2009a,
b,
2010a,
b).
Samar
and
Leyte
are
the
third
and
sixth
largest
islands
of
the
Philippines,
which
are
closely
adjacent
to
each
other
in
the
eastern
part
of
the
country.
The
first
record
of
the
cicadas
from
Samar
was
made
by
Haupt
(1924)
followed
by
Hayashi
(1987),
Beuk
(1999),
Duffels
and
Zaidi
(2000),
and
Lee
(2010a),
resulting
in
7
species
hitherto
recorded
from
Samar:
Cryptotympana
accipiter
(Walker),
Orientopsaltria
inermis
(Stal),
Orientopsaltria
fuliginosa
(Walker),
Champaka
nigra
(Distant),
Champaka
virescens
(Distant),
Huechys
parvula
Haupt,
and
Huechys
phaenicura
(Germar).
The
records
of
the
cicadas
from
Leyte
were
made
by
Endo
and
Hayashi
(1979),
Hayashi
(1987),
and
Duffels
and
Zaidi
(2000),
resulting
in
4
species
hitherto
recorded
from
Leyte:
C
accipiter,
0.
fuliginosa,
Dundubia
rufivena
Walker,
and
Huechys
sanguinea
(de
Geer).
In
the
combined
fauna
of
Samar
and
Leyte,
9
species
of
5
genera
have
been
recorded
so
far.
With
addition
of
a
new
species,
described
below,
from
Leyte
and
additional
three
new
records
to
Samar,
the
cicada
faunae
of
Samar
and
Leyte
now
comprise
13
species
belonging
to
7
genera
as
listed
below.
This
list
of
species
is
based
on
literature
that
specifically
mentioned
the
faunae
of
Samar
and
Leyte
Islands
and
specimens
from
the
Institut
Royal
des
Sciences
Naturelles
de
Belgique,
Brussels
(IRSNB).
The
synonymic
list
for
each
species
includes
only
the
references
that
specifically
mentioned
the
faunae
of
Samar
and
Leyte,
in
addition
to
the
original
description
with
the
type
locality
(n),
which
is
listed
as
given
in
the
original
publication.
Nomenclature
for
family,
subfamily,
and
tribal
classification
gen-
erally
follows
that
of
Lee
(2010b),
provided
that
the
tribal
placement
of
Oncotympana
Stal
follows
that
of
Lee
(2011).
Taxonomic
checklist
Family
Cicadidae
Subfamily
Cicadinae
Tribe
Cryptotympanini
Genus
Cryptotympana
Stal
Cryptotympana
Stal,
1861:
613.
Type
species:
Tettigonia
atrata
Fabricius.
Cryptotympana
accipiter
(Walker)
Fidicina
accipiter
Walker,
1850:
83
[11:
Philippine
Islands].
Cryptotympana
accipiter:
Hayashi,
1987:
1;
Lee,
2010b:
3.
Distribution.
Philippines
(S.
Luzon,
Samar,
and
Leyte).
*
Fax:
+1
860
465
9730.
Tribe
Cicadini
E-mail
address:
cicaderolee@gmail.com
.
Subtribe
Oncotympanina
1226-8615/$
-
see
front
matter
C
Korean
Society
of
Applied
Entomology,
Taiwan
Entomological
Society
and
Malaysian
Plant
Protection
Society,
2011.
Published
by
Elsevier
B.V.
All
rights
reserved.
doi:10.1016/j.aspen.2011.01.002
254
Y.J.
Lee
/
Journal
of
Asia-Pacific
Entomology
14
(2011)
253-257
Remarks.
Lee
(2011)
placed
Oncotympanina
within
Cicadini
for
the
Philippine
genera
Oncotympana
Stal
and
Neoncotympana
Lee.
Genus
Oncotympana
Stal
Pomponia
(Oncotympana)
Stal,
1870:
710.
Type
species:
Pomponia
(Oncotympana)
pallidiventris
Stal.
Oncotympana:
Distant,
1905:
60,
70.
Dokuma
Distant,
1905:
69.
Type
species:
Dundubia
nigristigma
Walker.
Remarks.
Lee
(2011)
placed
nine
species
in
Oncotympana:
0.
pallidiventris
Stal,
0.
grandis
Lee
(Lee,
2010b:
Luzon),
0.
viridicincta
Stal,
0.
nigristigma
(Walker),
0.
simonae
Lee
(Lee,
2010b:
Luzon),
0.
brevis
Lee
(Lee,
2010b:
Luzon),
0.
undata
Lee
(Lee,
2010b:
Luzon),
0.
averta
Lee
(Lee,
2011),
and
a
species
(Mindanao)
undescribed
but
misidentified
as
0.
viridicincta
by
Endo
and
Hayashi
(1979),
all
from
the
Philippines.
Oncotympana
pallidiventris
Stal
Pomponia
(Oncotympana)
pallidiventris
Stal,
1870:
710
[TL:
Philippines].
Material
examined.
1
male,
2
females,
C.
Samar,
8
km
S.E.
of
Bagacay,
primary
forest
road,
250
m,
11°48.025'N
125°14.610'E,
13
III
2009,
J.H.
Lourens
and
K.
Cerny
(IRSNB).
Distribution.
Philippines
(Samar).
Remarks.
This
species
has
been
reported
by
previous
authors
from
the
Philippines
without
further
details
about
the
specific
islands
but
is
newly
recorded
here
from
Samar.
Subtribe
Dundubiina
Genus
0rientopsaltria
Kato
0rientopsaltria
Kato,
1944:
6.
Type
species:
Dundubia
duarum
Walker.
0rientopsaltria
inermis
(Stal)
Cosmopsaltria
inermis
Stal,
1870:
708
[11:
Philippines].
0rientopsaltria
inermis:
Lee,
2010a:
22.
Material
examined.
3
males,
1
female,
W.E.
Samar,
4
km
S.E.
of
Bagacay,
primary
forest
road,
140
m,
11°48'N
125°15'E,
25
VI
2006,
J.H.
Lourens
(IRSNB);
2
females,
C.
Samar,
8
km
S.E.
of
Bagacay,
primary
forest
road,
250
m,
11°48.025'N
125°14.610'E,
13
III
2009,
J.H.
Lourens
and
K.
Cerny
(IRSNB).
Distribution.
Philippines
(Samar
and
Mindanao).
0rientopsaltria
fuliginosa
(Walker)
Dundubia
fuliginosa
Walker,
1850:
60
[TL:
Philippine
Islands].
0rientopsaltria
fuliginosa:
Duffels
and
Zaidi,
2000:
275;
Lee,
2010a:
24;
Lee,
2010b:
13.
Material
examined.
1
female,
C.
Samar,
8
km
S.E.
of
Bagacay,
primary
forest
road,
250
m,
11°48.025'N
125°14.610'E,
13
III
2009,
J.H.
Lourens
and
K.
Cerny
(IRSNB).
Distribution.
Philippines
(Luzon,
Mindoro,
Homonhon,
Samar,
Leyte,
Panay,
Negros,
Mindanao,
and
Basilan).
Genus
Dundubia
Amyot
and
Audinet-Serville
Dundubia
Amyot
and
Audinet-Serville,
1843:
470.
Type
species:
Tettigonia
vaginata
Fabricius.
Dundubia
vaginata
(Fabricius)
Tettigonia
vaginata
Fabricius,
1787:
266
[11:
Sumatra].
Material
examined.
1
male,
C.
Samar,
8
km
S.E.
of
Bagacay,
primary
forest
road,
250
m,
11°48.025'N
125°14.610'E,
13
III
2009,
J.H.
Lourens
and
K.
Cerny
(IRSNB);
4
males,
2
females,
same
data
except
26
III
2009
(IRSNB).
Distribution.
S.
China,
Philippines
(Luzon,
Sibuyan,
Samar,
Panay,
Palawan,
and
Mindanao),
Malay
Archipelago,
S.
Myanmar,
and
India.
Remarks.
This
species
is
newly
recorded
from
Samar.
Dundubia
rufivena
Walker
Dundubia
rufivena
Walker,
1850:
59
[TL:
Java];
Endo
and
Hayashi,
1979:
36.
Distribution.
Philippines
(Leyte),
Thailand,
Malaysia
(peninsular
Malaysia
and
Sabah),
Indonesia
(Sumatra,
Nias,
Java,
Sumbawa,
Kalimantan,
and
Sulawesi),
and
New
Guinea.
Genus
Champaka
Distant
Champaka
Distant,
1905:70.
Type
species:
Pomponia
viridimaculata
Distant.
Remarks.
This
genus
was
resurrected
and
redefined
by
Lee
(2010a).
Champaka
nigra
(Distant)
Cosmopsaltria
nigra
Distant,
1888:
292
[TL:
Philippine
Islands].
Platylomia
nigra:
Beuk,
1999:38.
Champaka
nigra:
Lee,
2010a:
25;
Lee,
2010b:
14.
Material
examined.
2
females,
C.
Samar,
8
km
S.E.
of
Bagacay,
primary
forest
road,
250
m,
11°48.025'N
125°14.610'E,
26
III
2009,
J.H.
Lourens
and
K.
Cerny
(IRSNB).
Distribution.
Philippines
(Luzon,
Sibuyan,
Samar,
and
Mindanao).
Champaka
virescens
(Distant)
Platylomia
virescens
Distant,
1
9
0
5
:
66
[11:
Sarawak,
Borneo;
Philippines];
Beuk,
1999:46.
Champaka
virescens:
Lee,
2010a:
25;
Lee,
2010b:
14.
Material
examined.
4
males,
6
females,
W.E.
Samar,
4
km
S.E.
of
Bagacay,
primary
forest
road,
140
m,
11°48'N
125°15'E,
25
VI
2006,
Y.J.
Lee
/
Journal
of
Asia-Pacific
Entomology
14
(2011)
253-257
255
40
0
A
B
C
D
Fig.
1.
Huechys
funebris
sp.
nov.,
holotype,
male,
Hilusig,
Leyte
(IRSNB).
(A)
dorsal
view.
(B)
ventral
view.
(C)
ventral view
of
male
pygofer.
(D)
lateral
view
of
male
pygofer.
J.H.
Lourens
(IRSNB);
2
females,
Leyte,
Hilusig,
W.
of
Mahaplag,
Mt.
Balocaue,
690
m,
10°40.742'N
124°55.270'E,
15-19
V
2007,
J.H.
Lourens
(IRSNB);
1
male,
5
females,
C.
Samar,
8
km
S.E.
of
Bagacay,
primary
forest
road,
250
m,
11°48.025'N
125°14.610'E,
13
III
2009,
J.H.
Lourens
and
K.
Cerny
(IRSNB);
1
male,
1
female,
same
data
except
26
III
2009
(IRSNB).
Distribution.
Philippines
(Luzon
(including
Camiguin
Island),
Negros,
Samar,
Leyte,
Mindanao,
and
Basilan)
and
Borneo.
Subfamily
Cicadettinae
Tribe
Huechysini
Genus
Huechys
Amyot
and
Audinet-Serville
Huechys
Amyot
and
Audinet-Serville,
1843:
464.
Type
species:
Cicada
sanguinea
de
Geer.
Huechys
parvula
Haupt
Huechys
(Huechys)
parvula
Haupt,
1924:
211
[TL:
Mindanao;
Catbalogan;
Samar];
Metcalf,
1963:
24.
Huechys
parvula:
Lee,
2010a:
25.
Distribution.
Philippines
(Samar
and
Mindanao).
Huechys
phaenicura
(Germar)
Cicada
phaenicura
Germar,
1834:
76
[TL:
India
orientali].
Huechys
(Huechys)
phaenicura:
Haupt,
1924:
212
(phoenicura
[sic]);
Metcalf,
1963:
24.
Huechys
phaenicura:
Lee,
2009a:
2636;
Lee,
2010b:
24.
Distribution.
Philippines
(Luzon,
Samar, Palawan,
and
Balabac
Island),
Malaysia,
Indonesia
(Java),
and
India.
Huechys
sanguinea
(de
Geer)
Cicada
sanguinea
de
Geer,
1773:
221,
pl.
33
(17)
[TL:
Chine].
Huechys
sanguinea:
Endo
and
Hayashi,
1979:
38;
Lee,
2009a:
2637.
Distribution.
S.
China
(including
Hainan),
Taiwan,
Philippines
(Leyte
and
Palawan),
Vietnam,
Thailand,
Malay
Peninsula,
Timor,
Borneo,
Indonesia
(Sumatra),
Myanmar,
and
India.
Huechys
funebris
sp.
nov.
(Fig.
1)
Type
material.
Holotype:
male
(Fig.
1),
"Coll.
I.R.Sc.N.B.
//
Philippines
fi
Philippines
Leyte
Hilusig
//
W.
of
Mahaplag
//
Mt.
Balocaue
600
m
/1
10°43'N
124°55'E
//
27-28
June
2006
//
JH
Lourens
leg"
(printed
yellow
and
white
label)
(IRSNB).
256
Y.
J.
Lee
/
Journal
of
Asia-Pacific
Entomology
14
(2011)
253-257
A
B
Fig.
2.
Nelcyndana
sp.,
female,
8
km
S.E.
of
Bagacay,
C.
Samar
(IRSNB).
(A)
dorsal
view.
(B)
ventral
view.
Etymology.
The
Latin
funebris
means
"of
a
funeral"
or
"black"
in
reference
to
the
entirely
black
abdominal
tergites
of
this
species,
which
are
not
seen
in
its
congeners.
Measurements
of
types.
Measurements
are
given
in
millimeter
for
1
male.
Length
of
body,
19.8;
width
of
head
including
eyes,
4.8;
width
of
pronotum,
6.2;
width
of
mesonotum,
4.9;
wing
span,
49.2.
Diagnosis.
This
species
is
closely
allied
to
Huechys
vidua
(White,
1846)
from
peninsular
Malaysia
as
this
species
has
a
relatively
narrow
head
and
pronotum,
a
relatively
long
male
abdomen,
black
abdominal
tergites,
and
9
forewing
apical
cells.
However,
this
species
differs
as
follows:
postclypeus
black
anteriorly
(reddish
ochraceous
anteriorly
in
H.
vidua);
male
abdominal
tergites
entirely
black
(with
small
red
areas
in
the
abdomen
of
H.
vidua);
hind
wing
anal
cell
2
(vannus)
broadly
margined
with
pale
fuscous
(narrowly
margined
in
H.
vidua).
The
entirely
black
abdominal
tergites
separate
this
species
from
all
other
species
of
Huechys.
Description
of
male
(Fig.
1).
Head
black.
Distance
between
lateral
ocelli
and
eyes
considerably
longer
than
the
distance
between
lateral
ocelli
but
much
narrower
than
twice
the
distance
between
lateral
ocelli.
Postclypeus
considerably
swollen.
Antenna
ochraceous
except
black
pedicel.
Ventral
part
of
head
black.
Postclypeus
reddish
orange
with
black
inverted-triangular
anteromedial
mark.
Anteclypeus,
rostrum,
lorum,
and
gena
entirely
black.
Pronotum
entirely
black
Mesonotum
black
with
a
pair
of
large
lateral
reddish
ochraceous
spots.
Cruciform
elevation
black
Thoracic
stemites
black
to
fuscous.
Legs
black
Fore-femur
with
primary
and
secondary
spines
and
two
small
subapical
spines:
primary
spine
largest,
acute,
obliquely
protruding
at
an
angle
of
about
50
degrees
to
femur; secondary
spine
about
two-thirds
as
long
as
primary
spine,
almost
upright;
subapical
spines
continuous
to
each
other,
about
one-third
as
long
as
secondary
spine.
Forewing
uniformly
blackish
brown
with
venation
mostly
black
but
apically
brown.
Basal
membrane
dark
gray.
Hind
wing
hyaline
tinged
with
brown,
margined
with
pale
fuscous,
and
with
venation
black
to
brown.
Hind
wing
anal
cell
2
(vannus)
broadly
margined
with
pale
fuscous.
Hind
wing
jugum
dark
gray.
Forewing
with
9
apical
cells.
Hind
wing
with
6
apical
cells.
Operculum
black;
longer
than
wide,
with
rounded
apical
margin;
not
reaching
posterior
margin
of
sternite
IL
Two
opercula
apart
from
each
other.
Abdominal
tergites
entirely
black.
Abdominal
stemites
reddish
orange
with
a
longitudinal
median
black
fascia.
Abdomen
longer
than
the
distance
from
head
to
cruciform
elevation.
Male
genitalia
(Fig.
1C
and
D):
Pygofer
oblong
in
ventral
view.
Dorsal
beak
nearly
equilateral
triangular.
Upper
lobe
of
pygofer
with
rounded
apex
in
lateral
view.
Anal
styles
about
as
large
as
median
lobe
of
uncus
in
lateral
view.
Median
lobe
of
uncus
slightly
elongated
semicircular
in
posterior
view.
Aedeagus
very
slender.
Thecal
pseudoparameres
present
dorsal
of
theca,
longer
than
theca.
Clasper
long,
slender,
curved
anterolaterad.
Distribution.
Philippines
(Leyte).
Tribe
Taphurini
Remarks.
Nelcyndana
Distant
was
erroneously
placed
in
Cicadettini
by
Lee
(2010a)
and
should
be
in
Taphurini
(cf.
Duffels,
2010).
Genus
Nelcyndana
Distant
Tibicen
(Nelcynda)
Stal,
1870:
716.
Nelcyndana
Distant,
1906:
139.
Type
species:
Tibicen
(Nelcynda)
tener
Nelcyndana
sp.
(Fig.
2)
Material
examined.
2
females,
C.
Samar,
8
km
S.E.
of
Bagacay,
primary
forest
road,
250
m,
11°48.025'N
125°14.610'E,
13
III
2009,
J.H.
Lourens
and
K.
Cerny
(IRSNB).
Distribution.
Philippines
(Samar).
Remarks.
This
species
is
believed
to
be
an
undescribed
new
species
and
thus
newly
recorded
from
Samar
but
could
not
be
described
due
to
unavailable
male
specimens.
The
female
specimens
have
mostly
light
green
head
and
thorax
(with
brown
pronotal
inner
area)
and
mostly
brown
abdomen
(Fig.
2).
Acknowledgments
I
am
indebted
to
Dr.
Jerome
Constant
(Institut
Royal
des
Sciences
Naturelles
de
Belgique,
Brussels)
for
the
loan
of
specimens.
I
am
grateful
to
anonymous
reviewers
for
suggestions
that
improved
the
manuscript.
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